- Rackham, Horsham District, West Sussex
- Racton, Chichester District, West Sussex
- Ringmer, Lewes District, East Sussex
- Ripe, Wealden District, East Sussex
- River Adur
- River Arun
- River Lavant
- River Ouse
- River Rother
- Robertsbridge, Rother District, East Sussex
- Rodmell, Lewes District, East Sussex
- Roffey, Horsham District, West Sussex
- Rogate, Chichester District, West Sussex
- Romney Marsh, Kent
- Rotherfield, Wealden District, East Sussex
- Rottingdean, The City of Brighton and Hove
- Rowfant, Crawley, West Sussex
- Rowhook, Horsham District, West Sussex
- Rudgwick, Horsham District, West Sussex
- Rumboldswyke, Chichester District, West Sussex
- Runcton, Chichester District, West Sussex
- Rushlake Green, Wealden District, East Sussex
- Rusper, Horsham District, West Sussex
- Rustington, Arun District, West Sussex
- Rye, Rother District, East Sussex
